Tutufa is a genus of sea snails, marine gastropod mollusks in the family Bursidae, the frog shells.[1] They have an Indo-West Pacific distribution.[2]
| Tutufa | |
|---|---|
| Two views of a shell of Tutufa bufo | |
| Five views of a shell of Tutufa bubo | |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Mollusca |
| Class: | Gastropoda |
| Subclass: | Caenogastropoda |
| Order: | Littorinimorpha |
| Superfamily: | Tonnoidea |
| Family: | Bursidae |
| Genus: | Tutufa Jousseaume, 1881 |
| Type species | |
| Murex rana bubo | |
| Synonyms[1] | |
| |
Description
editTutufa includes the largest bursids,[2] with the largest species, Tutufa bardeyi, attaining a shell height of up to 420 mm,[3] Tutufa bubo attaining a shell height of up to 300 mm,[3] and Tutufa tenuigranosa attaining similar sizes to T. bubo.[4] Other species, such as Tutufa bufo and Tutufa rubeta, are smaller.[3] Unlike most bursids, in which the varices are separated by 180° and form a pair of rows on opposite sides of the shell, the varices of Tutufa are separated by 240° and do not line up neatly, resembling cymatiids in this respect.[5][2] However, like all bursids and unlike cymatiids, Tutufa has an anal canal.[5] The shell is turreted. The whorls are nodose. The siphonal canal is very short and recurved.[6]
Taxonomy
editHistorically, the various species of Tutufa were considered to belong to one variable species. This species was known as Bursa lampas, Ranella lampas, or Triton lampas, using a species name coined by Carl Linnaeus that is now considered to apply to the triton shell Charonia lampas.[2] Linnaeus's original description of the species, which he called Murex lampas, referenced illustrations of shells of both taxa, but he stated that the species inhabited the Mediterranean, matching the range of Charonia lampas but not any of the species of Tutufa. In 1914, Edgar Albert Smith determined that the correct application of the name lampas was for the Mediterranean triton shell, not the bursid species, and recognized several varieties of the species that he called Bursa (Tutufa) rubeta.[7] The genus name Tutufa had originally been proposed in 1881 by Jousseaume with Murex lampas as the type species, but in 1977 the International Commission on Zoological Nomenclature ruled that since Jousseaume presumably meant the bursid species incorrectly known as Bursa lampas at the time, the type species of Tutufa should be Murex rana bubo, now known as Tutufa bubo, to match Jousseaume's intention.[2]
Species
editSpecies within the genus Tutufa include:
- Tutufa bardeyi (Jousseaume, 1881)[8]
- Tutufa boholica Beu, 1987[9]
- Tutufa bubo (Linnaeus, 1758)[10]
- Tutufa bufo (Röding, 1798)[11]
- Tutufa hitoshiikedai Thach, 2023
- Tutufa kevini Thach, 2023
- Tutufa lucsegersi Thach, 2023
- Tutufa nigrita Mühlhäusser & Blöcher, 1979[12]
- Tutufa oyamai (Habe, 1973)[13]
- Tutufa robusta T. Cossignani, 2009
- Tutufa rubeta (Linnaeus, 1758)[14]
- Tutufa tenuigranosa (E.A. Smith, 1914)[15]
- Tutufa truongi Thach, 2023
References
edit- 1 2 Tutufa Jousseaume, 1881. WoRMS (2010). Tutufa Jousseaume, 1881. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species on 19 June 2010 .
- 1 2 3 4 5 Beu, A. G. (1981-02-17). "Australian gastropods of the family Bursidae. Part 1. The families of Tonnacea, the genera of Bursidae, and revision of species previously assigned to Tutufa Jousseaume, 1881". Records of the Australian Museum. 33 (5): 248–324. doi:10.3853/j.0067-1975.33.1980.200. ISSN 0067-1975.
- 1 2 3 Beu, A.G.; Bouchet, P.; Tröndlé, J. (2012-05-31). "Tonnoidean gastropods of French Polynesia". Molluscan Research. 32 (2). doi:10.11646/mr.32.2.1. ISSN 1323-5818.
- ↑ Beu, A. G. (1986). "Taxonomy of gastropods of the families Ranellidae (= Cymatiidae) and Bursidae. Part 2. Descriptions of 14 new modern Indo‐West Pacific species and subspecies, with revisions of related taxa". New Zealand Journal of Zoology. 13 (3): 273–355. doi:10.1080/03014223.1986.10422668. ISSN 0301-4223.
- 1 2 Vokes, Emily H. (1973-04-11). "Notes on the fauna of the Chipola Formation – XIV. On the occurrence of Bursa (Mollusca: Gastropoda), with comments on the genus". Tulane Studies in Geology and Paleontology. 10 (2).
- ↑ Adams, H. & Adams, A. (1853-1858). The genera of Recent Mollusca; arranged according to their organization. London, van Voorst. Vol. 1: xl + 484 pp.; vol. 2: 661 pp.; vol. 3: 138 pls
- ↑ Smith, Edgar Albert (1914). "Note on Bursa (Tutufa) Rubeta (Bolten) = Triton lampas (Lamarck et auct.)". Journal of Conchology. 14 (8): 226–231. doi:10.5962/p.406161. ISSN 0022-0019.
- ↑ Tutufa bardeyi (Jousseaume, 1881). Retrieved through: World Register of Marine Species on 19 June 2010.
- ↑ Tutufa boholica Beu, 1987. Retrieved through: World Register of Marine Species on 19 June 2010.
- ↑ Tutufa bubo (Linnaeus, 1758). Retrieved through: World Register of Marine Species on 19 June 2010.
- ↑ Tutufa bufo (Röding, 1798). Retrieved through: World Register of Marine Species on 19 June 2010.
- ↑ Tutufa nigrita Mühlhäusser & Blöcher, 1979. Retrieved through: World Register of Marine Species on 19 June 2010.
- ↑ Tutufa oyamai (Habe, 1973). Retrieved through: World Register of Marine Species on 19 June 2010.
- ↑ Tutufa rubeta (Linnaeus, 1758). Retrieved through: World Register of Marine Species on 19 June 2010.
- ↑ Tutufa tenuigranosa (E.A. Smith, 1914). Retrieved through: World Register of Marine Species on 19 June 2010.
- Thach, N. N. (2023). New shells of South Asia and Japan, Taiwan, China. Akron, Ohio: 48Hrs Book Company. 141 pp. [including 63 plates].